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He's referring to the Theory of Alien Propagation, I think.
And yes, the Aliens had natural predators. The comic is very explicit on that point.
Then again, the entire comic is theoretical, and not actually fact.
Correction -- there are two other comics in which the Aliens have natural predators, in fact:
- 'Aliens: Reapers'
- 'Aliens: Taste'
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